PARADISE VALLEY, Mont. — Long after the fanfare that surrounded the return of the gray wolf to Yellowstone National Park, the most telling chapter may have been written this year by an unruly pack of ...

Snow crunched underfoot as Mark Hebblewhite scanned the ridgelines of Canada’s Banff National Park. It was 1995, and the young biologist, fresh out of undergrad, was trailing one of the park’s most ...